What is the role of `urls.py` in handling 404 errors?

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What is the role of `urls.py` in handling 404 errors?
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`urls.py` is central to handling 404 errors. Django's URL resolver attempts to match the requested URL against the patterns defined in your `urls.py`. If no pattern matches, Django generates a 404 error. You can also define a custom handler for 404s by setting `handler404` in your root `urls.py`.